N. Vietnam, prov. Cao Bang, distr. Tra Linh, municipality Quoc Toan, vicinities of Lung Tao village (N 22°45′ E 106°19′), about 12 km from Cao Bang town to the NNE. Primary semideciduous open broadleaved forest on steep slopes and bluffs of remnant limestone ridge at 600-800 m alt. Epilite creeping fern on open vertical cliff. Not common.
CBL 1233 14 Dec. 1998
International Botanical Expedition of the U.S.A. National Geographic Society (grant No 6300-98)
Collectors: N.T.Hiep, N.Q.Binh, P.K.Loc, N.X.Tam, Averyanov L.