S. Vietnam, Prov. Lam Dong, distr. Lac Duong, municipalite Da Chay, 31 km to NE from Dalat city (N 12° 08′, E 108° 39′). Closed primary evergreen broadleaved mt forest on NNW macroslope
of Bi Dup mt system at 1600-1700 m alt. Terrestrial fern in shadow steep stream canyon slope. Rare. VH 4510 02.05.1997 International Botanical Expedition of the U.S.A. National Geographic Society (grant No 5803-96)
Collectors: L. Averyanov, N.Q. Binh, N.T. Hiep, P.K. Loc, P. Lowry